This article cites 14 articles, 4 of which can be accessed free P<P Published online 23 May 2012 in advance of the print journal. This article is free to access Subject collections (69 articles)molecular biology (55 articles)health and disease and epidemiology (537 articles)ecology Articles on similar topics can be found in the following collections Email alerting service hereright-hand corner of the article or click Receive free email alerts when new articles cite this article- sign up in the box at the top publication. Citations to Advance online articles must include the digital object identifier (DOIs) and date of initial online articles are citable and establish publication priority; they are indexed by PubMed from initial pub...