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Abbreviations used in this paper: ANOVA, analysis of variance; AP, acceptor peptide; CRD, cysteine-rich domain; diHcRed, divalent Heteractis crispa red; FLIM, fluorescence lifetime imaging microscopy; FRET, Förster resonance energy transfer; IP, immunoprecipitate; LBD, ligand-binding domain; PY, phosphotyro-sine; SA, streptavidin; WB, Western blot; Wt, wild type