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The Natural Resource Publication series addresses natural resource topics that are of interest and applicability to a broad readership in the National Park Service and to others in the management of natural resources, including the scientific community, the public, and the NPS conservation and environmental constituencies. Manuscripts are peer- reviewed to ensure that the information is scientifically credible, technically accurate, appropriately written for the intended audience, and is designed and published in a professional manner. Natural Resource Reports are the designated medium for disseminating high priority, current natural resource management information with managerial application. The series targets a general, diverse audience, and may contain NPS policy considerations or address sensitive issues of management applicability. Examples of the diverse array of reports published in this series include vital signs monitoring plans; "how to " resource management papers; proceedings of resource management workshops or conferences; annual reports of resource programs or divisions of the Natural Resource Program Center; resource action plans; fact sheets; and regularly- published newsletters. Views and conclusions in this report are those of the authors and do not necessarily reflect policies of the National Park Service. Mention of trade names or commercial products does not constitute endorsement or recommendation for use by the National Park Service. Printed copies of reports in these series may be produced in a limited quantity and they are only available as long as the supply lasts. This report is also available from the Geologi