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Conservation of beneficial arthropods in agriculturallandscapes is increasingly justified based on the value of the services they provide to society. The annual value of natural enemies and native pollinators to agriculture has recently been estimated at nearly $8 billion in the US alone (Losey and Vaughn 2006). These arthropod-medi-ated ecosystem services (AMES) include biological con-trol of insect pests, worth $4.5 billion, and pollination of crops, worth $3.1 billion, each year. Despite the impres-sive economic value of these services, there is widespread concern over both the current and future status of benefi-cial arthropods. Farm intensification, urbanization, habi-tat fragmentation, climate change, diseases, and pesti-cides all threaten the services provided to agriculture by these arthropods (Allen-Wardell et al. 1998; Kremen et al. 2002; NRC 2007). Recognition of this situation has stim