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Keywords: Cryptic species tergreen group of Gaultheria (Angiospermae: Ericaceae: Gaultherieae) were estimated from separate and the Pacific Rim. The group is strongly supported as monophyletic, with the presence of methyl salicylate and a fleshy calyx surround-ing the capsule as likely synapomorphies (Bush et al., 2009a). Diplycosia and Tepuia are supported as monophyletic and nest as early-diverging clades within Gaultheria (Bush et al., 2009a). Phylogenetic results have only partly corroborated previous 2005). They occur from subtropical rainforests (400 m elevation) to alpine meadows (to 4500 m elevation), can be trees up to ca. 3 m tall to creeping or procumbent shrublets (ca. 3 cm tall), and are variable in leaf anatomy, inflorescence, bracteole position, fruit color, and seed morphology (see, e.g., Fritsch et al., 2008). The Sino-Himalayan Mountains and Malesia are the primary and secondary centers of diversity for Gaultheria in East Asia, respectively, each possessing more species of this genus than any other area of com-parable size in the western Pacific Rim (i.e., from Japan to India an