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tence within landscapes. Little attention has been paid to the impact of the distribution of the fragments trasting outcomes of fragmentation are related to how much origi-nal forest remains after fragmentation, the size and isolation of individual fragments and the distribution of the fragments over the landscape (Laurance et al., 2002; Pardini et al., 2010; Prugh et al., 2008). For example, lowland forests in Jamaica were cleared ts on forest struc-nce et al. al forests species co tion and diversity (Baldeck et al., 2013; Garcia-Lopez et al. Harms et al., 2001; John et al., 2007; Laurance et al., 2010), ing that the distribution of the fragments itself will, for a large part, determine the type and diversity of taxa preserved across the landscape (Virgos, 2001). Since forest fragmentation is generally a non-random process, with people targeting areas that are easily accessible or have good soil properties for agricultural production, forest fragments are usually restricted to areas located on steep slopes, poor soils or with inaccessible topography (Summerville and Crist, 2004; Vellend et al., 2008). The spatial distribution o