How maladaptation can structure olutionary island

underemphasizes an important and fundamental aspect of evolutionary ecology. Namely, the degree of local adapta-tion varies on a continuum from highly adapted to poorly adapted, with some degree of maladaptation being com-mon in nature [17]. On the one hand, simply recognizing the existence of Opinionhave seen an accumulation of research supporting this idea, both in the field of community genetics (see Glossary) [11,12] and more recently in the field of eco-evolutionary dynamics, the latter focusing explicitly on how rapid evo-lution can influence populations, communities, and ecosys-tems on contemporary timescales [13–16]. Much current research in eco-evolutionary dynamics focuses on scenarios where natural selection causes local adaptation to divergent habitats (Figure 1A) and thereb