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Establishing the primary axes (anterior-posterior and dorsal-ventral) is one of the first steps in patterning bilateral animals. In Drosophila, this process is well understood at the molecular level. One of the molecules that have been shown to be absolutely critical in patterning wild type embryos is the homeoprotein, Bicoid (Bcd) (St Johnston et al., 1989). Loss of bicoid (bcd) function results in embryos that lack all anterior structures, including the head, thorax, and some anterior abdominal segments (Frohnhofer and Nusslein-Volhard, 1986; St Johnston, 1995). However, the overall polarity of the remaining abdominal segments is retained. The bcd message is deposited into the egg by the mother, and factors binding to its 3’UTR localize it to the anterior pole of the egg (Berleth et al., 1988), via a migration along a polarized array of microtubules (Gonzalez-Reyes et al., 1995; Schnorrer et al., 2000). Translation of the bcd mRNA generates an anterior-to