RESEARCH ARTICLE Surface metrics: an alternative to patch metrics for the quantification of landscape structure

Abstract Modern landscape ecology is based on the patch mosaic paradigm, in which landscapes are conceptualized and analyzed as mosaics of discrete patches. While this model has been widely success-ful, there are many situations where it is more meaningful to model landscape structure based on continuous rather than discrete spatial heterogeneity. The growing field of surface metrology offers a variety of surface metrics for quantifying landscape gradients, yet these metrics are largely unknown and/ or unused by landscape ecologists. In this paper, we describe a suite of surface metrics with potential for landscape ecological application. We assessed the redundancy among metrics and sought to find groups of similarly behaved metrics by examining metric performance across 264 sample landscapes in western Turkey. For comparative purposes and to evaluate the robustness of the observed patterns, we examined 16 different patch mosaic models and 18 different landscape gradient models of landscape structure. Surface metrics were highly redundant, but less so than patch metrics, and consistently aggregated into four cohesive clusters of similarly behaved metrics representing surface roughness, shape of the surface height distribution, and angular and radial surface texture. While the surface roughness metrics have strong analogs among the patch metrics, the other surface components are largely unique to landscape gradients. We contend that the surface properties we identified are nearly universal and have potential to offer new insights into landscape pattern–process relationships