ICHAEL

, 41–49. The basal relationships of the hyperdiverse insect order Coleoptera (beetles) have proven difficult to resolve. Examination of beetle suborder relationships using 18S ribosomal DNA reveals a previously unproposed relationship among the four major lineages: [(Archostemata(Myxophaga(Adephaga, Polyphaga)))]. Adding representatives of most other insect orders results in a non-monophyletic Coleoptera. However, constraining Coleoptera and its suborders to be monophyletic, in analyses of beetle and outgroup sequences, also results in the above beetle relationships, with the root placed between Archostemata and the remaining suborders