This type occurs principally on moraines composed of well-to-moderately-well-drained heterogeneous soils with inherently high moisture and nutrient availability. Vegetation Description Structurally, these uneven-aged forests were characterized by large volumes of coarse, woody debris lying beneath multi-storied canopies of different-aged cohorts, with supercanopies composed of trees centuries old (Tyrell and Crow 1994). The dominant tree species are among the most moisture and nutrient-demanding species in the eastern U.S.A., and their distribution is confined to glacial landforms underlain by fertile soils (Woods 2000, Whitney 1986). Composition of the ground flora and understory varies along a moisture-nutrient gradient, and typically con...